He was an immunologist who did pioneering work on allergies. He developed the concept of a daily pollen count for the benefit of hay-fever sufferers.
He was a doctor in the British army in Singapore during WWII, and spent three-and-a-half years in Japanese POW camps.
He was made an MBE in 2015 for services to allergy research.
He studied medicine at the University of Oxford, and worked at St Mary's Hospital in Paddington, London. He worked with Sir Alexander Fleming, who discovered penicillin.