As Hook and Cora arrive in Storybrooke, Regina struggles to gain the trust of the town, as flashbacks show Snow and Charming disagree over how to handle Regina now that they have taken over the kingdom.
Gold's plans to cross the town line to find his son without losing his memories are thwarted by Hook, forcing Belle to intervene, as flashbacks show her befriend Mulan on a quest to destroy a beast.
As a stranger poses a potential threat to Storybrooke, Cora approaches Regina, and Whale battles his inner demons, as flashbacks detail information about Victor Frankenstein's life.
Emma and Henry accompany Gold on his quest to find his son, while Mary Margaret, David and Leroy deal with a vengeful giant, as flashbacks detail information about his life.
Emma is shocked when she learns the true identity of Gold's son, as flashbacks show when Rumplestiltskin was called to war. Back in Storybrooke, Regina, Cora and Hook search for Rumplestiltskin's dagger.
Hook hunts down Gold in New York and Mary Margaret learns of Regina and Cora's plan for the dagger, as flashbacks show a young Snow White faced with a difficult choice when her mother falls ill.
Mary Margaret is tempted by dark magic and David, Emma and Neal attempt to protect Gold from Regina and Cora, as flashbacks show a young Cora seek revenge against the royal family after they degrade her in public.
Emma and David enlist Gold's help to protect Mary Margaret from a vengeful Regina, as flashbacks show a father and son stumble upon Storybrooke when the curse was first cast.
Mary Margaret encounters a completely wooden August and Neal's fiancée arrives in Storybrooke, as flashbacks show August seek the help of a healer in Phuket when he starts to turn into wood.
Emma grows suspicious of Tamara and Hook helps Regina retrieve the fail-safe key that will destroy Storybrooke, as flashbacks show the Evil Queen go undercover in order to kill Snow White.
As Greg and Tamara set out to destroy Storybrooke, the inhabitants of the town race to stop the destruction, while flashbacks show Hook take Baelfire on as a crew member on his ship.
Tension runs high as Hook's ship sets sail for Neverland to rescue Henry, Neal travels to Rumplestiltskin's castle with Mulan, and Henry finds himself on the run from the Lost Boys after Greg and Tamara are ambushed.
Emma meets Peter Pan, who gives her a map that will reveal Henry's whereabouts as soon as she accepts her true identity, while flashbacks show the Evil Queen present Snow with an offer, which Charming does not want her to take.
To find Pan's hideout, Hook suggests tracking down Tinker Bell, who has a bitter past with Regina, and Robin Hood and Mulan help Neal get to Neverland.
Gold is reunited with Neal, as flashbacks show Rumplestiltskin go in search of a missing Baelfire, who has followed a mysterious figure who steals away children with his music.
Mary Margaret, Emma, David and Hook attempt to rescue Neal and Gold and Regina begrudgingly agree to work together to take down Pan, as flashbacks show Snow White help Ariel get acquainted with Prince Eric.
Gold and Regina send Ariel to Storybrooke to help Belle locate an artefact that could help take down Pan, while Emma, Neal and Hook attempt to find Dark Hollow in order to capture Pan's shadow.
As Pan leads Henry to a cave, Emma and the others race to save him, as flashbacks show when a young Rumplestiltskin was given a magical item that could help him make a fresh start with his father.
As Henry's life hangs in the balance, the race is on to stop Pan from gaining full power from the heart of the truest believer, as flashbacks show when Regina first adopted Henry.
As Hook's ship returns to Storybrooke, a well-hidden Pan puts a new plan into motion, while flashbacks show Snow and Charming go in search of a mythical being that could stop the Evil Queen cold in her tracks.